Publications, talks and data

Publications, talks and data

Publications resulting from the PRIDE programme:


Discover the unique mollusk fauna of the Black Sea - Identication sheets in four languages to download.

Deliverable D4.3 Outreach programme established and approved (2016), by Frank Wesselingh, Aleksandre Gogaladze, Caroline van Impelen, Niels Raes.


Sands, A.F, Glöer, P, Gürlek, M.E, Albrecht, C, & Neubauer, T.A. (2020). A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution96, 25–66. doi:10.3897/zse.96.48312

Sands, A.; Sereda, S.; Stelbrink, B.; Neubauer, T.A.; Lazarev, S.; Wilke, T. & Albrecht, C. (accepted): Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography.

Wesselingh, F.P.; Neubauer, T.A.; Anistratenko, V.V.; Vinarski, M.V., Yanina, T.; ter Poorten, J.J.; Kijashko, P.; Albrecht, C.; Anistratenko, O.Y.; D’Hont, A.; Frolov, P.; Gándara, A.M.; Gittenberger, A.; Gogaladze, A.; Karpinsky, M.; Lattuada, M.; Popa, L.; Sands, A.F.; van de Velde, S.; Vandendorpe, J. & Wilke, T. (2019): Mollusc species from the Pontocaspian region – an expert opinion list. ZooKeys, 827, 31-124.

Sands, A.F.; Apanaskevich, D.A.; Matthee, S.; Horak, I.G. & Matthee, C.A. (2017): The effect of host vicariance and parasite life history on the dispersal of the multi‐host ectoparasite, Hyalomma truncatum. Journal of Biogeography, 44, 1124-1136.

 Old lake versus young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus) Arthur F. Sands, Thomas A. Neubauer, Saeid Nasibi, Majid Fasihi Harandi, Vitaliy V. Anistratenko, Thomas Wilke and Christian Albrecht, Royal Society Open Science, September 2019.

Past and future impact of the winter North Atlantic Oscillation in the Caspian Sea catchment areaSri D. Nandini-Weiss, Matthias Prange, Klaus Arpe, Ute Merkel, Michael Schulz. International Journal of Climatology, 14 October 2019,
Amplitude, frequency and drivers of Caspian Sea lake‐level variations during the Early Pleistocene and their impact on a protected wave‐dominated coastline. Elisabeth L. Jorissen,  Hemmo A. Abels,  Frank P. Wesselingh,  Sergei Lazarev,  Vusala Aghayeva &  Wout Krijgsman, Journal of Sedimentology.
Magneto-biostratigraphic age constraints on the palaeoenvironmental evolution of the South Caspian basin during the Early-Middle Pleistocene (Kura basin, Azerbaijan)
by Sergei Lazarev, Elisabeth L. Jorissen, Sabrina van de Velde, Lea Rausch, Marius Stoica, Frank  P. Wesselingh, Christiaan G. C. Van Baak, Tamara  A. Yanina, Elmira Aliyeva, Wout Krijgsman, Quaternary Science Reviews, Volume 222, 15 October 2019, 105895. doi: 10.1016/j.quascirev.2019.105895

An Early Pleistocene anomalohaline water ostracod fauna from lake deposits of the Homo Erectus-bearing Kocabaş locality (SW Turkey). Lea Rausch & Marius Stoica, Acta Palaeontologica Romaniae (2019), 

Historical faunal exchange between the Pontocaspian Basin and North AmericaJustine Vandendorpe, Christiaan G. C. van Baak, Björn Stelbrink, Diana Delicado, Christian Albrecht and Thomas Wilke, Ecology and Evolution (2019),

Social network analysis and the implications for Pontocaspian biodiversity conservation in Romania and Ukraine: A comparative study, Aleksandre Gogaladze, Niels Raes, Koos Biesmeijer, Camelia Ionescu, Bianca Pavel, Mikhail O. Son, Natalia Gozak, Vitaliy Anistratenko and Frank P. Wesselingh. doi:

A conservation palaeobiological approach to assess faunal response of threatened biota under natural and anthropogenic environmental changeSabrina van de Velde, Elisabeth L. Jorissen, Thomas A. Neubauer, Silviu Radan, Ana Bianca Pavel, Marius Stoica, Christiaan G. C. Van Baak, Alberto Martínez Gándara, Luis Popa, Henko de Stigter, Hemmo A. Abels, Wout Krijgsman, and Frank P. Wesselingh, Biogeosciences, 16, 2423–2442 (2019),

Contributions of biogeographical functions to species accumulation may change over time in refugial regionsArthur F. Sands, Sergej V. Sereda, Björn Stelbrink, Thomas A. Neubauer, Sergei Lazarev, Thomas Wilke and Christian Albrecht
Journal of Biogeography 46:1274–1286 (2019).

The Late Pleistocene mollusk fauna of Selitrennoye (Astrakhan province, Russia): A natural baseline for endemic Caspian Sea faunasSabrina van de Velde, Tamara A.Yanin, Thomas A.Neubauer and Frank P. Wesselingh, J. Great Lakes Res. 1–13, 2019

Where should we draw the lines between dinocyst “species”? Morphological continua in Black Sea dinocysts, Thomas M. Hoyle, Manuel Sala-Pérez, and Francesca Sangiorgi, J. Micropalaeontol., 38, 55–65, 2019.

Differential impact of anthropogenic pressures on Caspian Sea ecoregions, Matteo Lattuada, Christian Albrecht and Thomas Wilke, Marine Pollution Bulletin.

Mollusc species from the Pontocaspian region – an expert opinion list, Frank P. Wesselingh, Thomas A. Neubauer, Vitaliy V. Anistratenko, Maxim V. Vinarski, Tamara Yanina, Jan Johan ter Poorten, Pavel Kijashko, Christian Albrecht, Olga Yu. Anistratenko, Anouk D’Hont, Pavel Frolov, AlbertoMartínez Gándara, Arjan Gittenberger, Aleksandre Gogaladze, Mikhail Karpinsky, Matteo Lattuada, Luis Popa, Arthur F. Sands, Sabrina van de Velde, Justine Vandendorpe, Thomas Wilke.

Drivers of dominance shifts between invasive Ponto-Caspian dreissenids Dreissena polymorpha (Pallas, 1771) and Dreissena rostriformis bugensis (Andrusov, 1897), Anouk D’Hont, Adriaan Gittenberger, A. Jan Hendriks and Rob S.E.W. Leuven, Aquatic Invasions, volume 13, issue 4, November 2018.

Quaternary time scales for the Pontocaspian domain: Interbasinal connectivity and faunal evolution, W. Krijgsman, A. Tesakov, T. Yanina, S. Lazarev, G. Danukalova, C.G.C. Van Baak, J. Agustí, M.C. Alçiçek, E. Aliyeva, D. Bista, A. Bruch, Y. Büyükmeriç, M. Bukhsianidze, R. Flecker, P. Frolov, T.M. Hoyle, E.L. Jorissen, U. Kirscher, S.A. Koriche, S.B. Kroonenberg, D. Lordkipanidze, O. Oms, L. Rausch, J. Singarayer, M. Stoica, S. van de Velde, V.V. Titov, F.P. Wesselingh. Elsevier, Earth-Science Reviews, Volume 188, January 2019, Pages 1-40.

Sedimentary architecture and depositional controls of a Pliocene river-dominated delta in the semi-isolated Dacian Basin, Black Sea, Elisabeth L. Jorissen, Arjan de Leeuw, Christiaan G.C. van Baak , Oleg Mandic, Marius Stoica, Hemmo A. Abels, Wout Krijgsman. Elsevier, Sedimentary Geology, Volume 368, June 2018, Pages 1-23.

Scientific drilling projects in ancient lakes: Integrating geological and biological histories, Thomas Wilke, Bernd Wagner, Bert Van Bocxlaer, Christian Albrecht, Daniel Ariztegui, Diana Delicado, Alexander Francke, Mathias Harzhauser, Torsten Hauffe, Jens Holtvoeth, Janna Just, Melanie J. Leng, Zlatko Levkov, Kirsty Penkman, Laura Sadori, Alister Skinner, Björn Stelbrink, Hendrik Vogel, Frank Wesselingh, Thomas Wonik, Global and Planetary Change, 143, Elsevier, 2016.

Articles in press

The rise and demise of mollusk species in the Pontocaspian region – a request for help, Arthur Sands and Justine Vandendorpe, Spirula Quarterly, 406, Netherlands, 2016.


Sedimentary architecture of a Pliocene river-dominated delta flowing into the semi-isolated Dacian Basin (Romania). Elisabeth L. Jorissen, Arjan de Leeuw, Christiaan G.C. van Baak, Oleg Mandic, Marius Stoica, Hemmo A. Abels, Wout Krijgsman ; oral presentation at conference ''International Meeting of Sedimentologists'' in Toulouse, France, October  2017.

Plio-Pleistocene stratigraphy and environmental evolution in the Ponto-Caspian region, Thomas Hoyle, RCMNS meeting in Athens (15th Congress of the RCMNS, Exploring a physical laboratyory: The Mediterranean Basin), on the 6th September 2017.
PRIDE: Understanding Longterm Natural Biotic and Abiotic Drivers of Lake System Change in the Pontocaspian Basin, Christian Albrecht, Frank Wesselingh, Thomas Wilke & the PRIDE team, Abstract book of conference “Speciation in Ancient Lakes 7”, SIAL7,  August 17th – 20th, University of Windsor, Canada, 2015.
Posters 2017
Past & Future E-P evolution in the Caspian Catchment: Climate Model, Lake Model & Observations, Sri Nandini, American Geophysical Union Conference 2017 in New Orleans, USA in December 11th-15th, 2017. 

Past and future impact of North Atlantic teleconnection patterns on the hydroclimate of the Caspian catchment area in CESM1.2.2 and observations, by Sri Nandini, M. Prange, M. Schulz, Geophysical Research Abstracts Vol. 19, EGU2017-4665 European Geoscience Union (EGU) general assembly, 2017.

Historical Climate Change Impacts on the Hydrological Processes of the Ponto-Caspian Basin, by Sifan A. Koriche, Joy S. Singarayer, Michael T. Coe, Sri Nandini, Matthias Prange, Hannah Cloke, and Dan Lunt, Geophysical Research Abstracts Vol. 19, EGU2017-1313 European Geoscience Union (EGU) general assembly, 2017.

Testing the connectivity history of the Black Sea and the Caspian Sea during the Pleistocene, using strontium isotopes, by D. Bista, R. Flecker, D.A. Richards, Goldschmidt conference 2017, Paris.

Testing the connectivity history of the Black Sea and the Caspian Sea during the Pleistocene, using strontium isotopes, by D. Bista, R. Flecker, D.A. Richards, UK Paleoclimate Society Conference (PACS) 2017, Cardiff.

Late-Sarmatian S.L. – Maeotian Littoral-Deltaic Environments of the Dacian Basin along the Slănicul de Buzau section: Facies Analysis and Paleogeography, Lazarev S.Yu. , de Leeuw A., Stoica M., Mandic O., van Baak C., Krijgsman W. Book of Abstracts of the 15th Congress of the RCMNS, Athens, Greece, 2017.

Dinoflagellates from the Pontocaspian region: from biodiversity to phylogeny, Manuel Sala-Pérez et al., 2017, at DINO11 – 11th International Conference on Modern and Fossil Dinoflagellates - July 17 to 21 Boudreaux, France.  

Transarctic Interchange, Justine Vandendorpe, at the Conference: "Pontocaspia: Where East meets West" in Azov, Russia, August 2017.

Differential Impact of Anthropogenic Pressures on Caspian Sea Ecoregions, Matteo Lattuada, poster at the 10th Conference of the Giessen Graduate centre for Life sciences (GGL), Giessen, Germany and at the Conference: "Pontocaspia: Where East meets West" in Azov, Russia, August 2017.

Drivers of the progradation of a river-dominated delta in a semi-isolated basin: example of the Dacian Basin (Romania). E. L. Jorissen, A. de Leeuw, C. G. C. van Baak, O. Mandic, M. Stoica, H. A. Abels, W. Krijgsman; poster at conference "Pontocaspia: Where East meets West" in Azov, Russia, August 2017.

Dinoflagellates from the Pontocaspian region: from biodiversity to phylogeny, Manuel Sala-Pérez et al., 2017, poster at conference "Pontocaspia: Where East meets West" in Azov, Russia, August 2017.

THE EARLY PLEISTOCENE AGE MODEL OF THE KURA BASIN ALONG THE GOYCHAY AND HAJIGABUL SECTIONS: PALEOMAGNETIC APPROACH FOR DATING AND CORRELATION, S. Lazarev, E.L. Jorissen, C.G.C. van Baak, W. Krijgsman, poster at conference "Pontocaspia: Where East meets West" in Azov, Russia, August 2017.

Evolutionary Events and Diversity in Pontocaspian Mollusk Groups by Combined Molecular and Fossil Data, Justine Vandendorpe, at EU PRIDE mid-term review meeting in Leiden, 16 February 2017.

Posters and talks in 2015 and 2016

Climate Change Impact on Hydrological Processes of the Ponto-Caspian Basin, by S. Koriche, J. Singarayer, D. Lunt, S. Nandini, M. Prange, at the INQUA ECR 2016 conference in Reading. (Best poster award).

Reconstructing the hydrological evolution of the Black Sea and the Caspian Sea during the last 2 Ma using strontium isotopes, by D. Bista, R. Flecker, D.A. Richards, at the INQUA ECR 2016 conference in Reading. (Second best poster award).

Mean Climate Variability in the Ponto-Caspian Basin in the past 150 years with CESM1.2.2, by S. Nandini, at the International Union for Quaternary Research (INQUA) Conference, 2016.

Holocene biodiversity changes in Pontocaspian planktonic communities, Manuel Sala-Perez, Suzanne A.G. Leroy, Anne E. Lockyer, International Conference on Ecological Sciences 2016, Sfecologie2016, 24-28 October in Marseille, 2016.

Trans-arctic Interchange, Justine Vandendorpe, at the 9th Conference of the Giessen Graduate centre for Life sciences (GGL), Giessen, Germany, 2016.

Modelling the distributions of bivalve species using spatial ecological parameters derived from observational data that predate the remote sensing era, Matteo Lattuada, Niels Raes, at the 9th Conference of the Giessen Graduate centre for Life sciences (GGL), Giessen, Germany, 2016.

Human footprint and biodiversity in the Ponto-caspian region, Matteo Lattuada, seminar talk at the University of Giessen, 2016.

Preliminary morphometric study of Hypanis plicata shells of the Razelm lake, Alberto M. GÁNDARA, Mihaela Isabela VADANA, Larisa-Bianca DUMITRESCU, Oana Paula POPA, Luis Ovidiu POPA, The 7th International Zoological Congress of “Grigore Antipa” Museum 18-21 November 2015, Bucharest – Romania.

The Razim-Sinoe lake complex in the Danube Delta, Romania: a geological refuge for the endangered Pontocaspian fauna? Elisabeth L. Jorissen, Sabrina van de Velde, Gülçin Aygün, Yavar Moshifar, Hemmo A. Abels, Christiaan G.C. van Baak, Wout Krijgsman, Suzanne A.G. Leroy, Silviu Radan, Marius Stoica, Frank P. Wesselingh, and the PRIDE-ITN team; poster at conference ''Nederlands Aardwetenschappelijk Congres'' in Eindhoven, Netherlands, April  2016.


All data and insights of the PRIDE programme will be publicly available through mostly Open Access publications and sharing our data. Taxonomic data will be organized and presented PC-TAX (the PontoCaspian Taxonomy platform). 

In the course of 2017 another online platform will become available, named >PC-IS (PontoCaspian Information System). The PC-IS will be a vehicle to combine climate and lake basin scenarios with biodiversity change in past and future and allow stakeholders and other interested people to explore the intricate relationships and roles of various drivers of biodiversity change in the Pontocaspian area. 


Outreach is central in the PRIDE programme. From 2015-2018 the Early Stage Researchers (ESRs) will investigate the stakeholders that should have an interest in understanding the drivers of the Pontocaspian biodiversity crisis and devise ways to engage them with our insights and results.

The various stakeholders will include regular academic institutes and researchers, students governmental organizations (from ministries to National Park management in the various Pontocaspian countries), NGO’s, journalists, but also local people as well the general public in the region and in the wider EU.

The ESRs will investigate the stakeholders and develop a policy how to reach out in their first year and become responsible for the execution of the outreach programme, together with their host institutes and aided by our partners in the last two years of their assignments. We intend to develop a far-ranging outreach strategy that will enable to link with very different target groups. Many hands will do the work.

This is the link to the Outreach programme that was established and approved in November 2016: Outreach programme